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Aquificota Wiki2Web Clarity Challenge

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Study Guide: The Aquificota Phylum: Characteristics, Taxonomy, and Evolutionary Insights

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The Aquificota Phylum: Characteristics, Taxonomy, and Evolutionary Insights Study Guide

Fundamental Characteristics and Habitat

The *Aquificota* phylum is primarily composed of Archaea that inhabit extreme environments.

Answer: False

Explanation: The *Aquificota* phylum consists of true bacteria, not Archaea, despite both groups often inhabiting extreme environments. They belong to the domain Bacteria.

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The name *Aquificota* translates to 'water maker' and is derived from the genus *Aquifex*, reflecting its ability to produce water through hydrogen oxidation.

Answer: True

Explanation: The name *Aquificota* is indeed derived from the genus *Aquifex*, meaning 'water maker,' which accurately describes its capacity for water production via hydrogen oxidation.

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*Aquificota* bacteria are typically found in temperate aquatic environments, avoiding harsh conditions.

Answer: False

Explanation: *Aquificota* bacteria are typically found in harsh aquatic environments, such as hot springs and deep-sea vents, rather than temperate conditions.

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*Aquificota* bacteria are characterized as Gram-positive, spore-forming cocci.

Answer: False

Explanation: *Aquificota* bacteria are characterized as Gram-negative, non-spore-forming rods, not Gram-positive, spore-forming cocci.

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Despite inhabiting similar extreme environments, *Aquificota* are classified as true bacteria, distinct from Archaea.

Answer: True

Explanation: Although *Aquificota* and Archaea can both inhabit extreme environments, *Aquificota* are classified as true bacteria within the domain Bacteria, distinguishing them from Archaea.

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*Aquificota* belong to the domain Eukaryota.

Answer: False

Explanation: *Aquificota* belong to the domain Bacteria, not Eukaryota.

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Being 'Gram-negative' means *Aquificota* bacteria have a thick peptidoglycan layer that retains crystal violet stain.

Answer: False

Explanation: Being 'Gram-negative' means *Aquificota* bacteria possess a cell wall structure that does *not* retain the crystal violet stain, due to their outer membrane and thinner peptidoglycan layer.

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*Aquificota* are known for producing highly resistant endospores to survive harsh conditions.

Answer: False

Explanation: *Aquificota* are characterized as non-spore-forming, meaning they do not produce highly resistant endospores.

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What is the fundamental nature of the *Aquificota* phylum?

Answer: A diverse group of bacteria known for inhabiting harsh environmental conditions.

Explanation: The *Aquificota* phylum represents a diverse group of true bacteria that are well-adapted to and found in harsh environmental conditions.

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From which genus does the name *Aquificota* originate, and what does it signify?

Answer: *Aquifex*, meaning 'water maker'.

Explanation: The name *Aquificota* originates from the genus *Aquifex*, which translates to 'water maker,' reflecting its ability to produce water through hydrogen oxidation.

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Which of the following is a typical habitat for *Aquificota* bacteria?

Answer: Harsh aquatic environments like springs and oceans.

Explanation: *Aquificota* bacteria are typically found in harsh aquatic environments, such as hot springs, geothermal pools, and deep-sea hydrothermal vents.

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What are the key morphological and cellular characteristics of *Aquificota* bacteria?

Answer: Gram-negative, non-spore-forming rods.

Explanation: *Aquificota* bacteria are characterized as Gram-negative, non-spore-forming rods.

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How are *Aquificota* distinguished from Archaea, despite often inhabiting similar extreme environments?

Answer: *Aquificota* are classified as true bacteria, belonging to the domain Bacteria.

Explanation: *Aquificota* are classified as true bacteria within the domain Bacteria, which fundamentally distinguishes them from Archaea, despite their shared preference for extreme environments.

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What is the domain classification of *Aquificota*?

Answer: Bacteria.

Explanation: *Aquificota* are classified under the domain Bacteria.

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What does 'Gram-negative' signify for *Aquificota* bacteria?

Answer: They possess a cell wall structure that does not retain the crystal violet stain.

Explanation: For *Aquificota* bacteria, being 'Gram-negative' means their cell wall structure does not retain the crystal violet stain during the Gram staining procedure.

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What is the significance of *Aquificota* being 'non-spore-forming'?

Answer: They do not produce dormant, resistant structures like endospores.

Explanation: The characteristic of *Aquificota* being 'non-spore-forming' signifies that these bacteria do not produce dormant, resistant structures such as endospores.

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Taxonomic Hierarchy and Nomenclature

The phylum *Aquificota* was validly published by Cavalier-Smith in 2020.

Answer: False

Explanation: The phylum *Aquificota* was validly published by Reysenbach in 2021, not by Cavalier-Smith in 2020.

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The *Aquificota* phylum currently includes over 100 genera and species.

Answer: False

Explanation: The *Aquificota* phylum currently encompasses 15 genera and 42 species, not over 100 genera and species.

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The three main classes comprising the *Aquificota* phylum are Aquificia, Desulfurobacteriia, and Thermosulfidibacteria.

Answer: True

Explanation: The *Aquificota* phylum is indeed composed of three distinct classes: Aquificia, Desulfurobacteriia, and Thermosulfidibacteria.

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The order Aquificales includes the families Aquificaceae and Desulfurobacteriaceae.

Answer: False

Explanation: The order Aquificales includes the families Aquificaceae and Hydrogenothermaceae, not Desulfurobacteriaceae. Desulfurobacteriaceae belongs to the Desulfurobacteriales order.

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The Desulfurobacteriales order is unique in containing only one family, the Desulfurobacteriaceae.

Answer: True

Explanation: The Desulfurobacteriales order is indeed characterized by containing only one family, the Desulfurobacteriaceae.

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*Thermosulfidibacter takaii* is not assigned to a specific family within *Aquificota* due to its phylogenetic distinctness from both Aquificales and Desulfurobacteriales.

Answer: True

Explanation: *Thermosulfidibacter takaii* is phylogenetically distinct from both the Aquificales and Desulfurobacteriales orders, leading to its non-assignment to a specific family within *Aquificota*.

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'Aquificaeota' (Oren et al. 2015) is a currently accepted name for the *Aquificota* phylum.

Answer: False

Explanation: 'Aquificaeota' (Oren et al. 2015) is listed as a historical synonym for the *Aquificota* phylum, implying it is not the currently accepted name.

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The currently accepted taxonomy of *Aquificota* relies on information from the LPSN and NCBI databases.

Answer: True

Explanation: The currently accepted taxonomy for *Aquificota* is indeed based on information from the List of Prokaryotic names with Standing in Nomenclature (LPSN) and the National Center for Biotechnology Information (NCBI).

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According to the LTP_10_2024 phylogenetic tree, the Aquificales order includes the Desulfurobacteriaceae family.

Answer: True

Explanation: The LTP_10_2024 phylogenetic tree indicates that the Aquificales order includes the Desulfurobacteriaceae family, among others.

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The GTDB 09-RS220 classification places *Aquificota* into three main classes: Desulfurobacteriia, Aquificia, and Thermosulfidibacteria.

Answer: False

Explanation: The GTDB 09-RS220 classification places *Aquificota* into two main classes: Desulfurobacteriia and Aquificia, not three.

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The kingdom classification for *Aquificota* is Pseudomonadati.

Answer: True

Explanation: According to the provided scientific classification, the kingdom for *Aquificota* is Pseudomonadati.

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The Genome Taxonomy Database (GTDB) uses 16S rRNA gene sequences for its standardized bacterial and archaeal taxonomy.

Answer: False

Explanation: The Genome Taxonomy Database (GTDB) bases its standardized bacterial and archaeal taxonomy on 120 marker proteins, not primarily on 16S rRNA gene sequences.

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The All-Species Living Tree Project (LTP) primarily bases its phylogenetic tree on 16S rRNA gene sequences.

Answer: True

Explanation: The All-Species Living Tree Project (LTP) primarily constructs its phylogenetic tree of prokaryotic species based on 16S rRNA gene sequences.

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The Aquificales order, according to the infobox, includes the family Desulfurobacteriaceae.

Answer: False

Explanation: According to the infobox, the Aquificales order includes Aquificaceae and Hydrogenothermaceae, while Desulfurobacteriaceae is associated with the Desulfurobacteriales order.

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The Desulfurobacteriales order is associated with the family Desulfurobacteriaceae in the infobox.

Answer: True

Explanation: The infobox indeed associates the Desulfurobacteriales order with the family Desulfurobacteriaceae.

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The kingdom Pseudomonadati is a specific, well-detailed grouping for *Aquificota* with many shared characteristics explicitly described in the source.

Answer: False

Explanation: While *Aquificota* is classified under the kingdom Pseudomonadati, the source material does not provide extensive details or shared characteristics for this grouping.

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Who is credited with the valid publication of the phylum *Aquificota* in 2021?

Answer: Reysenbach.

Explanation: Reysenbach is credited with the valid publication of the phylum *Aquificota* in 2021.

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How many genera and species are currently recognized within the *Aquificota* phylum?

Answer: 15 genera and 42 species.

Explanation: The *Aquificota* phylum currently encompasses 15 genera and 42 validly published species.

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Which of the following are the three main classes that comprise the *Aquificota* phylum?

Answer: Aquificia, Desulfurobacteriia, and Thermosulfidibacteria.

Explanation: The three main classes comprising the *Aquificota* phylum are Aquificia, Desulfurobacteriia, and Thermosulfidibacteria.

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Which families belong to the order Aquificales within the *Aquificota* phylum?

Answer: Aquificaceae and Hydrogenothermaceae.

Explanation: The order Aquificales includes the families Aquificaceae and Hydrogenothermaceae.

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What is the sole family recognized within the Desulfurobacteriales order?

Answer: Desulfurobacteriaceae.

Explanation: The Desulfurobacteriales order contains only one recognized family, the Desulfurobacteriaceae.

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Why is *Thermosulfidibacter takaii* not assigned to a specific family within the *Aquificota* phylum?

Answer: It is phylogenetically distinct from both Aquificales and Desulfurobacteriales orders.

Explanation: *Thermosulfidibacter takaii* is not assigned to a specific family within *Aquificota* due to its phylogenetic distinctness from both the Aquificales and Desulfurobacteriales orders.

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Which of the following is NOT a historical synonym for the *Aquificota* phylum?

Answer: 'Proteobacteria' (Garrity et al. 2005).

Explanation: 'Proteobacteria' is a distinct phylum and not a historical synonym for *Aquificota*.

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Which two primary databases are used for the currently accepted taxonomy of *Aquificota*?

Answer: LPSN and NCBI.

Explanation: The currently accepted taxonomy of *Aquificota* relies on information from the LPSN (List of Prokaryotic names with Standing in Nomenclature) and NCBI (National Center for Biotechnology Information) databases.

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According to the 16S rRNA based LTP_10_2024 phylogenetic tree, which family is associated with the Thermosulfidibacterales order within *Aquificota*?

Answer: Thermosulfidibacteraceae.

Explanation: The 16S rRNA based LTP_10_2024 phylogenetic tree associates the Thermosulfidibacterales order with the Thermosulfidibacteraceae family within *Aquificota*.

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Based on the 120 marker proteins from GTDB 09-RS220, what are the two main classes within the *Aquificota* phylum?

Answer: Desulfurobacteriia and Aquificia.

Explanation: Based on the 120 marker proteins from GTDB 09-RS220, the two main classes within the *Aquificota* phylum are Desulfurobacteriia and Aquificia.

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What kingdom is *Aquificota* classified under?

Answer: Pseudomonadati.

Explanation: The *Aquificota* phylum is classified under the kingdom Pseudomonadati.

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According to the infobox, which families are included in the Aquificales order?

Answer: Aquificaceae and Hydrogenothermaceae.

Explanation: According to the infobox, the Aquificales order includes the families Aquificaceae and Hydrogenothermaceae.

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Molecular Phylogeny and Markers

CSIs are used to differentiate between the Aquificales and Desulfurobacteriales orders.

Answer: True

Explanation: Conserved Signature Indels (CSIs) are indeed utilized to differentiate between the Aquificales and Desulfurobacteriales orders, as specific CSIs are unique to each group.

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*Aquificota* and Thermotogales bacteria share a unique 51-amino-acid insertion in their SecA preprotein translocase.

Answer: True

Explanation: Both *Aquificota* and Thermotogales bacteria share a unique 51-amino-acid insertion in their SecA preprotein translocase, serving as a distinct molecular signature.

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In 16S rRNA gene trees, *Aquificota* species typically branch far from Thermotogota, indicating a distant evolutionary relationship.

Answer: False

Explanation: In 16S rRNA gene trees, *Aquificota* species typically branch in close proximity to Thermotogota, suggesting a close evolutionary relationship, not a distant one.

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Phylogenetic studies based on Hsp70 and Hsp60 proteins strongly support a deep branching position for *Aquificota* and a close relationship with Thermotogota.

Answer: False

Explanation: Phylogenetic studies based on Hsp70 and Hsp60 proteins, along with other CSIs, do *not* support a deep branching position for *Aquificota* or a close relationship with Thermotogota; instead, they suggest a relationship with Proteobacteria.

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A three-amino-acid CSI in inorganic pyrophosphatase is unique to *Aquificota* and Proteobacteria, supporting their relationship.

Answer: False

Explanation: A *two*-amino-acid CSI in inorganic pyrophosphatase is uniquely shared by *Aquificota* and Proteobacteria, not a three-amino-acid CSI.

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Comparative genomic studies are crucial for identifying CSIs and understanding the evolutionary relationships within *Aquificota*.

Answer: True

Explanation: Comparative genomic studies are indeed essential for identifying conserved signature indels (CSIs) and for understanding the complex evolutionary relationships within the *Aquificota* phylum.

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A two-amino-acid CSI in inorganic pyrophosphatase is shared uniquely by *Aquificota* and Proteobacteria.

Answer: True

Explanation: A unique two-amino-acid conserved signature indel (CSI) in the inorganic pyrophosphatase protein is indeed shared by species from both *Aquificota* and Proteobacteria, supporting their evolutionary relationship.

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What are Conserved Signature Indels (CSIs) primarily used for in the context of *Aquificota*?

Answer: To serve as molecular markers indicating shared evolutionary history and distinguishing features.

Explanation: Conserved Signature Indels (CSIs) are primarily used as molecular markers to indicate shared evolutionary history and distinguishing features among organisms, including *Aquificota*.

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How do CSIs help differentiate between the Aquificales and Desulfurobacteriales orders?

Answer: By showing specific insertions or deletions in proteins unique to each order.

Explanation: CSIs help differentiate between the Aquificales and Desulfurobacteriales orders by revealing specific insertions or deletions in proteins that are unique to each group.

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What unique molecular signature is shared by *Aquificota* and Thermotogales bacteria?

Answer: A 51-amino-acid insertion in their SecA preprotein translocase.

Explanation: *Aquificota* and Thermotogales bacteria share a unique 51-amino-acid insertion in their SecA preprotein translocase.

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Where do *Aquificota* species typically branch in 16S rRNA gene trees?

Answer: In close proximity to the phylum Thermotogota, near the archaeal-bacterial branch point.

Explanation: In 16S rRNA gene trees, *Aquificota* species typically branch in close proximity to the phylum Thermotogota, often near the archaeal-bacterial branch point.

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What evidence contradicts a deep branching position for *Aquificota* and a close relationship with Thermotogota?

Answer: Phylogenetic studies based on other gene/protein sequences and CSIs in universal proteins.

Explanation: Phylogenetic studies utilizing other gene/protein sequences and conserved signature indels (CSIs) in universal proteins contradict a deep branching position for *Aquificota* and a close relationship with Thermotogota.

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What specific molecular evidence supports a relationship between *Aquificota* and Proteobacteria?

Answer: A unique two-amino-acid CSI in the inorganic pyrophosphatase protein.

Explanation: A unique two-amino-acid CSI in the inorganic pyrophosphatase protein provides specific molecular evidence supporting a relationship between *Aquificota* and Proteobacteria.

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Physiological Adaptations and Metabolism

As autotrophs, *Aquificota* bacteria are primary carbon fixers, converting inorganic carbon into organic compounds.

Answer: True

Explanation: *Aquificota* bacteria are autotrophs and play a vital ecological role as primary carbon fixers, converting inorganic carbon into organic compounds.

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Members of the Desulfurobacteriales are microaerophilic and can oxidize various compounds like sulfur.

Answer: False

Explanation: Members of the Desulfurobacteriales are strict anaerobes and exclusively oxidize hydrogen, whereas Aquificales are microaerophilic and can oxidize sulfur or thiosulfate.

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The SecA insertion contributes to thermostability by forming a network of water molecules that stabilize ADP/ATP binding at high temperatures.

Answer: True

Explanation: The 51-amino-acid insertion in SecA contributes to thermostability by forming a network of water molecules near the ADP/ATP binding site, which stabilizes ADP/ATP binding at high temperatures.

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The high G+C content in *Aquificota* rRNAs is essential for maintaining secondary structure stability at high growth temperatures.

Answer: True

Explanation: The very high G+C content in *Aquificota* rRNAs is indeed crucial for maintaining the stability of their secondary structures, which is essential for functionality at their high growth temperatures.

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*Aquificota* and Thermotogota are both classified as psychrophilic eubacteria, thriving in cold environments.

Answer: False

Explanation: *Aquificota* and Thermotogota are both classified as thermophilic eubacteria, meaning they thrive in high-temperature environments, not psychrophilic (cold-loving) environments.

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A 'microaerophile' requires high concentrations of oxygen to survive.

Answer: False

Explanation: A 'microaerophile' requires oxygen, but only at lower concentrations than those present in the atmosphere, not high concentrations.

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*Aquificota* are considered hyperthermophilic organisms, thriving in extremely hot environments.

Answer: True

Explanation: *Aquificota* are classified as thermophilic eubacteria, and in some contexts, are considered hyperthermophilic, thriving in extremely hot environments.

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What is the ecological role of *Aquificota* bacteria in their environments?

Answer: They are primary carbon fixers, converting inorganic carbon into organic compounds.

Explanation: *Aquificota* bacteria are autotrophs and serve as primary carbon fixers, playing a crucial role in converting inorganic carbon into organic compounds.

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What is a key physiological difference between members of the Desulfurobacteriales and Aquificales orders?

Answer: Desulfurobacteriales exclusively oxidize hydrogen, while Aquificales can oxidize sulfur or thiosulfate in addition to hydrogen.

Explanation: A key physiological difference is that Desulfurobacteriales are strict anaerobes that exclusively oxidize hydrogen, whereas Aquificales are microaerophilic and can oxidize various compounds, including sulfur or thiosulfate, in addition to hydrogen.

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How does the 51-amino-acid insertion in SecA contribute to the thermostability of *Aquificota* bacteria?

Answer: It stabilizes ADP/ATP binding at high temperatures by forming a water molecule network.

Explanation: The 51-amino-acid insertion in SecA contributes to thermostability by forming a network of water molecules that stabilize ADP/ATP binding at high temperatures.

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What is the significance of the high G+C content in the rRNAs of *Aquificota*?

Answer: It is crucial for the stability of their secondary structures at high growth temperatures.

Explanation: The high G+C content in *Aquificota* rRNAs is crucial for maintaining the stability of their secondary structures at the high growth temperatures they inhabit.

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What type of eubacteria are *Aquificota* and Thermotogota classified as, based on their temperature preference?

Answer: Thermophilic.

Explanation: Both *Aquificota* and Thermotogota are classified as thermophilic eubacteria, thriving in high-temperature environments.

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What is a 'microaerophile' in the context of Aquificales?

Answer: An organism that requires oxygen, but only at lower concentrations than atmospheric levels.

Explanation: In the context of Aquificales, a 'microaerophile' is an organism that requires oxygen for survival, but only at concentrations lower than those present in the atmosphere.

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Evolutionary Insights and Gene Transfer

The 51-amino-acid insertion in SecA in *Aquificota* and Thermotogales is believed to have developed due to lateral gene transfer.

Answer: False

Explanation: Phylogenetic studies indicate that the shared 51-amino-acid insertion in SecA in *Aquificota* and Thermotogales developed independently due to selective pressure from high temperatures, not through lateral gene transfer.

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CSIs in proteins like Hsp70 and AlaRS suggest a specific relationship between *Aquificota* and the phylum Proteobacteria.

Answer: True

Explanation: Evidence from CSIs in proteins such as Hsp70 and AlaRS strongly supports an evolutionary relationship between *Aquificota* and the phylum Proteobacteria, particularly Campylobacterota.

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The frequent grouping of *Aquificota* with Campylobacterota is primarily due to their identical 16S rRNA gene sequences.

Answer: False

Explanation: The frequent grouping of *Aquificota* with Campylobacterota is explained by some authors as a result of frequent horizontal gene transfer between these groups, not identical 16S rRNA gene sequences.

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Lateral gene transfer was the primary mechanism for the shared SecA insertion between *Aquificota* and Thermotogales.

Answer: False

Explanation: Phylogenetic studies suggest that the shared SecA insertion between *Aquificota* and Thermotogales developed independently due to selective pressure, not primarily through lateral gene transfer.

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Selective pressure from cold environments led to the SecA insertion in *Aquificota*.

Answer: False

Explanation: Selective pressure from *high-temperature* environments, not cold environments, is believed to have led to the SecA insertion in *Aquificota*, as this feature enhances thermostability.

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Informational genes are generally more prone to horizontal gene transfer than non-informational genes.

Answer: False

Explanation: Informational genes, involved in core cellular processes, are generally considered *less* prone to horizontal gene transfer compared to non-informational genes.

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The placement of *Aquificota* near the archaeal-bacterial branch point in 16S rRNA trees suggests a recent evolutionary origin.

Answer: False

Explanation: The placement of *Aquificota* near the archaeal-bacterial branch point in 16S rRNA trees suggests they might represent one of the *earliest diverging* lineages within the Bacteria, indicating an ancient, not recent, evolutionary origin.

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Cavalier-Smith suggested that *Aquificota* are closely related to Archaea.

Answer: False

Explanation: Cavalier-Smith suggested that *Aquificota* are closely related to the Proteobacteria, not Archaea.

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What is the proposed reason for the shared 51-amino-acid insertion in SecA in *Aquificota* and Thermotogales?

Answer: Independent development in response to selective pressure from high temperatures.

Explanation: The shared 51-amino-acid insertion in SecA in *Aquificota* and Thermotogales is proposed to have developed independently due to selective pressure from high-temperature environments, indicating convergent evolution.

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Which bacterial phylum is suggested to have a specific relationship with *Aquificota* based on CSIs in important proteins?

Answer: Proteobacteria.

Explanation: Evidence from CSIs in important proteins suggests a specific evolutionary relationship between *Aquificota* and the phylum Proteobacteria.

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How do some authors explain the frequent grouping of *Aquificota* with Campylobacterota?

Answer: As a result of frequent horizontal gene transfer between these groups.

Explanation: Some authors explain the frequent grouping of *Aquificota* with Campylobacterota as a result of frequent horizontal gene transfer, likely due to shared ecological niches.

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What is 'lateral gene transfer' (LGT)?

Answer: The movement of genetic material between organisms other than by parent-offspring transmission.

Explanation: Lateral gene transfer (LGT) is defined as the movement of genetic material between organisms by means other than vertical transmission from parent to offspring.

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What is the significance of the 'archaeal-bacterial branch point' in *Aquificota* phylogeny?

Answer: It suggests *Aquificota* might represent one of the earliest diverging lineages within the Bacteria.

Explanation: The placement of *Aquificota* near the archaeal-bacterial branch point in 16S rRNA trees suggests they might represent one of the earliest diverging lineages within the Bacteria, indicating an ancient evolutionary origin.

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